As is usual with mysterious "permission denied" errors, SELinux was the underlying cause. ...
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ID Status Resolution Sev Product Comp Assignee Reporter Summary
36986 NEW --- nor Sisyphus virtualb sin zxwarior Рассинхронизация runlevel в сервисах vboxadd{,-service}

File a new bug in the "virtualbox-guest-utils" component of the "Sisyphus" product